to·ti·pal·mate (t??t?-p?l?m?t?)adj. Having webbing that connects all four toes, as in waterbirds such as pelicans, gannets, and boobies.[Latin t?tus, whole; see total + palmate.]to?ti·pal·ma?tion n.American Heritage® Dictionary of the English Language, Fifth Edition. Copyright © 2016 by Houghton Mifflin Harcourt Publishing Company. Published by Houghton Mifflin Harcourt Publishing Company. All rights reserved.totipalmationthe state of having all four toes fully webbed, as water birds. ? totipalmate, adj.See also: Birds-Ologies & -Isms. Copyright 2008 The Gale Group, Inc. All rights reserved.