to·ti·pal·mate (t??t?-p?l?m?t?)adj. Having webbing that connects all four toes, as in waterbirds such as pelicans, gannets, and boobies.[Latin t?tus, whole; see total + palmate.]to?ti·pal·ma?tion n.American Heritage® Dictionary of the English Language, Fifth Edition. Copyright © 2016 by Houghton Mifflin Harcourt Publishing Company. Published by Houghton Mifflin Harcourt Publishing Company. All rights reserved.totipalmate (?t??t??pælm?t; -?me?t) adj (Zoology) (of certain birds) having all four toes webbed[C19: from Latin t?tus entire + palmate, from Latin palm?tus shaped like a hand, from palma palm1] ?totipal?mation nCollins English Dictionary ? Complete and Unabridged, 12th Edition 2014 © HarperCollins Publishers 1991, 1994, 1998, 2000, 2003, 2006, 2007, 2009, 2011, 2014